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After defeating the various Floor Masters of the six prior levels, Hakuno Kishinami and Saber arrive at the top floor of the Moon Cell. Their opponent will be Leonardo B. Harwey—the strongest Master in the history of the Holy Grail War and the current fan favorite, following his previous victory. As they reach a field of flowers among floating isles, Hakuno, Saber, and Rin Toosaka come across a hooded man tending to the blossoms. Saber immediately draws her blade before the cloaked figure, who reveals himself as Prince Gawain: the Platinum Saber and Knight of the Round Table. However, when Gawain insists that he has no interest in fighting, the four discuss the infinite possibilities for the Holy Grail, with Hakuno determined to return the Moon Cell to its original form. Their discussion ends with a clangor from afar, ringing through the skies. The threadbare Gawain, knowing its significance, informs the three that not much time remains for their lives. He beckons them towards the final battleground, where the two Sabers shall duel once more to determine the future of mankind. [Written by MAL Rewrite]

A technological hell masquerading as paradise, Tsukimihara Academy is an artificial high school that serves as the setting for the next Holy Grail War. Created by the Moon Cell computer, the school is inhabited by Earth-projected souls who have even the slightest aptitude for being a "Master." Of these 256 souls, 128 will be chosen for the main tournament and granted a Servant. With all of the Masters selected, the Academy activates a purge, targeting the remaining lifeforms for elimination. Awakening in a pool of his own blood, Hakuno Kishinami refuses to die. Fueled by unknown feelings of hatred, he vows to fight for survival. As he struggles to escape from a relentless pursuer, he finds a crimson blade plunged into the ground; and by pulling it out, Hakuno summons his own Servant, Saber, who instantly destroys his pursuer in a flurry of rose petals. With his newfound power, Hakuno must now begin his journey to Moon Cell's core, the Angelica Cage. There, he will unveil the reason for this artificial world and the secrets of his own blood-soaked past. [Written by MAL Rewrite]

Fifty years ago, mysterious creatures known as the "Huge" suddenly appeared all over the world, posing an existential threat to humanity. To defend against these menaces, soldiers in the form of teenage girls known as "Lilies" use the energy "Magie" within their bodies to power weapons called "Counter Huge ARMS" or "CHARMS." After being saved by a Lily two years ago while evacuating from a Huge attack, Riri Hitotsuyanagi vowed to become a Lily herself. She enrolls in the prestigious academy of Yurigaoka, a training ground for Lilies, and discovers that one of her classmates is Yuyu Shirai—the same Lily who rescued her. Much to her confusion, however, the person she thought to be cheerful turns out to be quite antisocial and prefers to fight alone. Even so, Riri still desires to get along with Yuyu and is willing to do anything she can to reach that goal. With Yuyu and the other Lilies by her side, Riri's journey on becoming one of them has just bloomed! [Written by MAL Rewrite]

Fifty years ago, mysterious creatures known as the "Huge" suddenly appeared all over the world, posing an existential threat to humanity. To defend against these menaces, soldiers in the form of teenage girls known as "Lilies" use the energy "Magie" within their bodies to power weapons called "Counter Huge ARMS" or "CHARMS." After being saved by a Lily two years ago while evacuating from a Huge attack, Riri Hitotsuyanagi vowed to become a Lily herself. She enrolls in the prestigious academy of Yurigaoka, a training ground for Lilies, and discovers that one of her classmates is Yuyu Shirai—the same Lily who rescued her. Much to her confusion, however, the person she thought to be cheerful turns out to be quite antisocial and prefers to fight alone. Even so, Riri still desires to get along with Yuyu and is willing to do anything she can to reach that goal. With Yuyu and the other Lilies by her side, Riri's journey on becoming one of them has just bloomed! [Written by MAL Rewrite]