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When one space colony declared its independence from the Earth Federation, the devastating 7th Space War, an all-out war between Earth and space, resulted. The Federation responded to the Space Revolutionary Army with mobile suits, called Gundams. However, the Space Revolutionary forces played their trump card and dropped hundreds of space colonies onto the Earth, plunging the planet into a seven-year-long nuclear winter. The Federation collapsed, but the Space Revolutionary Army was unable to invade the Earth in the aftermath of the colony drop. Fifteen years have passed. The year is now After War 0015, and a New Federation has sprung up on Earth to restore order. In space, the colonial leaders have been rebuilding their own forces as well. By chance, fifteen-year-old Garrod Ran has discovered an old Federation mobile suit, the Gundam X, and now he uses it to help out the Vulture ship Freeden in its struggle to keep the powers that be from repeating the mistakes of the past.

The United Earth Sphere Alliance is a powerful military organization that has ruled over Earth and space colonies with an iron fist for several decades. When the colonies proclaimed their opposition to this, their leader was assassinated. Now, in the year After Colony 195, bitter colonial rebels have launched "Operation Meteor," sending five powerful mobile suits to Earth for vengeance. Built out of virtually indestructible material called Gundanium Alloy, these "Gundams" begin an assault against the Alliance and its sub organization OZ. One Gundam, whose pilot has taken the name of the slain colony leader Heero Yuy, is forced to make a crash landing into the ocean after an atmospheric battle against OZ's ace pilot Zechs Marquise. Upon coming ashore, he is found by Relena Peacecraft, daughter of a peace-seeking politician, who witnesses Heero's descent to Earth. Although neither of them realize it yet, this encounter will have a profound impact on both their lives, as well as those on Earth and in space colonies. [Written by MAL Rewrite]

In the year Cosmic Era 0071, the space colony Heliopolis remains neutral in the great war raging across the galaxy between Coordinators, human beings whose biological traits have been altered before birth, and Naturals, unaltered people who remain on the planet Earth. The Naturals' deep hatred of the Coordinators drove the advanced beings into space, seeking shelter in man-made colonies. Kira Yamato is a Coordinator and university student on Heliopolis, when his life is thrown into disarray as ZAFT, the military organization composed of rebellious Coordinators, attacks the colony in an effort to steal a set of five state-of-the-art military mobile suits known as Gundams. While ZAFT manages to make off with four of the mobile suits, Kira take control of the final Gundam, the Strike. Surviving the battle, Kira and his college friends join the crew of the Archangel, a ship run by the Earth Alliance, and the young soldiers experience the horrors of war and the loss that comes with it. [Written by MAL Rewrite]

In the year Cosmic Era 0071, a wartime brawl between two mobile suits results in the destruction of a neutral country. Shinn Asuka finds his entire family killed as casualties of the violence, and swears his life toward a vengeful pursuit of the Earth's Natural forces, under the impression they were the ones responsible for his family's death. Shinn works his way up through the ZAFT military forces, eventually gaining access to the cockpit of the prototype Impulse mobile suit. Athrun Zala, a veteran of the recent war employed in the service of diplomat Cagalli Yula Athha, finds himself drawn into a new conflict between the Naturals and the Coordinators: human beings artificially enhanced before birth and hated by the Naturals for their biological advantages. As the new war rears its head and violence becomes inevitable, Athrun finds himself as a mentor to the bitter Shinn. Athrun must steer the young pilot towards a mindset of mercy before his rage and desire for vengeance consumes him. [Written by MAL Rewrite]